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The USS Sephora is sent to investigate the Sulaco. 17 weeks later the Sephora arrives, and its marine crew begin investigating. While searching the ship Corporal Cristopher T. Winter, Private Peter O'Neal and Private Bella Clarison discover Weyland-Yutani PMCs commanding the Sulaco and using Sephora marines as hosts to the xenomorphs. The hostile corporate mercenaries attack the Sephora using the Sulaco's weaponry and both ships are destroyed after the Sephora retaliates. Winter, O'Neal, Bella, Captain Jeremy Cruz, Sephora android Bishop, and dropship pilot Lieutenant Lisa Reid crash-land on LV-426 and take shelter in the ruins of the Hadley's Hope colony complex. Captain Cruz orders all the other survivors from the Sephora to gather at the Hadley Hope's ruins. While traveling the ruins of the colony, mutated and radioactive xenomorphs that only react to sound and movement, and are prone to exploding violently due to their unstable chemical makeup are discovered, as well as the corpse of Sulaco marine William Hudson.

Private Bella is found to be infected with a xenomorph after a scan by Bishop. As a result, she is escorted by Winter and O'Neal to a Weyland-Yutani xenomorph research facility, set near the derelict extraterrestrial "Engineer" ship from Alien, in order to surgically remove the embryo. However, an interrogated Weyland-Yutani physician explains to them that the infection can't be treated since extracting the alien will also kill the host anyway. Bella dies when her parasite matures while she is saying her goodbyes to Winter and O'Neal.

Hicks is rescued from the PMCs. After gathering all the remaining Sephora Marines, Captain Cruz orders Hicks and all of his men to attack the Weyland-Yutani facility and seize the only remaining faster-than-light space cruiser left on the planet. While boarding the escaping ship, Captain Cruz sacrifices himself to kill an Alien Queen hidden inside the ship's cargo bay. Winter, O'Neal, Reid, Bishop and Hicks capture and kill Michael Weyland, who is revealed to also be a Bishop-model android, and not the real Weyland. Searching for military intelligence, Bishop is connected to "Weyland's" CPU and states that he has "everything", setting the stage for a war between the Colonial Marines and the Weyland-Yutani Corporation.